If
A + B means B is the brother of A;
A x B means B is the husband of A;
A - B means A is the mother of B and
A % B means A is the father of B,
which of the following relations shows that Q is the grandmother of T ?

A. Q - P + R % T
B. P x Q % R - T
C. P x Q % R + T
D. P + Q % R - T